E. Gordon Grau

PhD University of Delaware (Biology), 1978

Professor, Department of Zoology
Researcher, Hawai'i Institute of Marine Biology
Director, Hawai`i Sea Grant College Program

Hawai'i Institute of Marine Biology
P.O. Box 1346
Kane'ohe, Hawai'i 96744
phone: (808) 236-7406
FAX: (808) 236-7443

Environmental physiology and comparative endocrinology of estuarine fish

[publications] [graduate students]
The neuroendocrine system regulates development and physiology in fish and coordinates their interactions with their environment. Our studies are based on the idea that reproduction, development, immune function and environmental adaptation are regulated through the orderly release of hormones by the neuroendocrine system, which integrates information from genes and the environment. Such studies can improve the way we utilize and protect the marine organisms, the renewable resources of marine environments. Studies in developmental endocrinology can be used to increase the food supply through aquaculture without compromising environmental health. These studies can also provide new avenues for restoring populations of endangered species, for limiting noxious organisms and for clarifying the effects of endocrine-disrupting chemicals. These chemicals enter the environment through human activity. Their negative impact on marine animals, including fish, can be profound because hormones play a crucial role in controlling development.

The regulation of salt and water balance is a fundamental requirement of life. The structure and function of macromolecules depend closely on their interactions with water and its solutes. Few factors affect the distribution and evolution of an organism as extensively as osmoregulation. Thus, organisms invest considerable energy in controlling precisely the composition of both intracellular and extracellular fluids. In fish, osmoregulation typically consumes 25-50% of the total metabolic output, possibly the largest single component. Osmotic equilibrium is maintained only through the precise interplay of a major portion of the neuroendocrine array. Indeed, the maintenance of osmotic balance in seriously ill patients is among the most difficult challenges in medicine. In view of the cost and importance of osmoregulation, it may seem ironic that the mechanisms that monitor and regulate osmotic balance are so poorly understood. Closer attention reveals the impediment, the typically complex structure and arrangement of osmoreceptive cells and tissues. These problems are obviated by the use of the osmosensitive prolactin cell of a fish pituitary. Prolactin plays a fundamental role in freshwater osmoregulation, and prolactin cells are directly sensitive to extracellular osmolality. Our studies are aimed at elaborating the cellular mechanisms that mediate this osmoreceptive response. The information these studies provide has found useful application in such diverse areas as biotechnology, medicine and aquaculture.

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Representative publications

Shepherd BS, Sakamoto T, Hyodo S, Nishioka RS, Ball C, Bern HA, Grau EG. 1999. Is the primitive regulation of pituitary prolactin (tPRL177 and tPRL188) secretion and gene expression in the euryhaline tilapia (Oreochromis mossambicus) hypothalamic or environmental? J Endocrinol 161:121-129.

Morrey C, Nakamura M, Kobayashi T, Grau EG, Nagahama Y. 1998. P450scc-like immunoreactivity throughout gonadal restructuring in the protogynous hermaphrodite, Thalassoma duperrey. Int J Devel Biol 42:811-816.

Shepherd BS, Sakamoto T, Mori I, Nishioka RS, Richman NH III, Madsen S, Hirano T, Bern HA, Grau EG. 1997. Somatotropic actions of the homologous growth hormone (tGH) and prolactins in the euryhaline tilapia, Oreochromismossambicus. PNAS 94:2068-2072.

Weber GM, Powell JFF, Park M, Fisher WH, Rivier JE, Nanakorn U, Parhar IS, Grau EG, Sherwood NM. 1997. Primary structures for three gonadotropin-releasing hormone (GnRH) molecules and evidence that GnRH functions as a prolactin-releasing factor in tilapia. J Endocrinol 155:121-132.

Current students

Larry Riley (PhD)
endocrine regulation of growth and reproduction in tilapia
Andre Seale (PhD)
effects of cRFa on PRL and GH secretion in the tilapia
Jason Collier (MS)

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Last update: 5 December 2000
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